2017 | Affiliate Member, LMCB, UCL, London2007-2017 | Scientific Director, Max F Perutz Laboratories, Vienna1999-2007 | Professor of Cell Biology, Yale Medical School1988-1999 | Principal Scientist, ICRF (now CRUK), London1985-1988 | Professor and Chair of Biochemistry, Dundee1977-1985 | Group Leader, EMBL, Heidelberg1972-1977 | Post-doc NIMR, London & Biochemistry, Cambridge1969-1972 | PhD Biochemistry, Cambridge

2011 | Member, Austrian Academy of Sciences2007 | Eduard Buchner Prize, German Biochemical Society1999 | Fellow of the Royal Society, London1986 | Elected EMBO Member