Evans Lab

Susan Evans
Evans Lab News
Resources Paul Barrett BBSRC skull mechanics project
Arnau Bolet Marc Jones Ryoko Matsumoto Elizabeth Ward Jérémy Anquetin Marianne Pearson
Evans Lablab
Marc Jones, Susan Evans, Paul Barret, Jérémy Anquetin and Ryoko Matsumo

Research area:
Research in the Evans lab focuses on the morphological evolution of lissamphibians and non-avian reptiles and its implications for macroevolution, phylogenetic relationships, function, palaeoecology, and biogeography. Techniques and methods used include cladistic analysis, geometric morphometrics, Multibody Dynamics Analysis, Computed Tomography, Scanning Electron Microscopy, classical comparative anatomy, and tissue histology.

Current research projects:
We are currently involved in a number of UK and international research projects including a BBSRC funded project investigating skull mechanics in reptiles using Rigid Body computer models and Finite Element Analysis in collaboration with University of Hull and Hull-York Medical School.

Networking activities and society involvement:
Researchers in the Evans lab are active members of the Society of Vertebrate Paleontology; regularly attend LERN, CEE and Grant Museum events; contribute to the NHM-UCL palaeotetrapod journal club; and with the Goswami, Upchurch, and Chatterjee groups, organise the ADaPTIVE (Anatomy, Diversity and Phylogeny: Trends in Vertebrate Evolution) inter-lab meetings at UCL. 

Please note:
Research in the Evans Lab does not involve experimentation on live animals.

Currently our main source of funding comes from the Biotechnology and Biological Research Council (BBSRC). However, we have also received funding from the Leverhulme Trust, Natural Environment Research Council (NERC), Palaeontological Association (PalAss), Royal Society, British Council, UCL Graduate School, UCL Bogue, University of London Central Research fund, and National Geographic.

Current lab members:

Professor Susan E Evans - Lab head

Marianne Pearson - PhD student

Elizabeth Ward - PhD student

Dr Marc E H Jones - Research associate

Arnaut Bolet - Honorary research associate/PhD student

Dr Paul M Barrett - Honorary research associate

Dr David Whiteside - Honorary research associate

Dr Ryoko Matusmoto - Honorary research associate

Dr Jérémy Anquetin - Honorary research associate

Emma Humphries - Honorary research associate

Neil Brocklehurst - Honorary research associate

PhD project opportunities in vertebrate palaeontology and morphology:
Please contact Professor Susan E Evans or Dr Paul M Barrett.

Location (and postal address):
Evans Lab, Main Anatomy Building (Ground floor: G08)
Research Department of Cell and Developmental Biology
Gower Street, UCL,
University College London
London, UK

Nearest tube stations are Warren Street, Euston, Euston Square, and Goodge Street

Lab phone number:
+44 (0) 20 7679 0160 (x30160)

Shared fax number:
+44 (0) 20 7679 7349

Selected recent publications from members of the Evans lab


  • Evans SE, Groenke JR, Jones MEH, Turner AH, Krause DW. 2014. New material of Beelzebufo, a hyperossified frog (Amphibia: Anura) from the Late Cretaceous of Madagascar. PLoS ONE 9(1): e87236. DOI:10.1371/journal.pone.0087236
  • Anquetin J, Deschamps S, Claude J. 2014 The rediscovery and redescription of the holotype of the Late Jurassic turtle Plesiochelys etalloni. PeerJ 2:e258. DOI:10.7717/peerj.258


  • Bolet A, Evans SE. 2013. Lizards and amphisbaenians (Reptilia, Squamata) from the late Eocene of Sossís (Catalonia, Spain), Palaeontologia Electronica 16(1): 8A 23p palaeo-electronica.org/content/2013/412-eocene-squamates-from-spain
  • Gröning F, Jones MEH, Curtis N, Herrel A, O'Higgins P, Evans SE, Fagan MJ. 2013. The importance of accurate muscle modelling for biomechanical analyses: a case study with a lizard skull. Journal of Royal Society Interface 10: DOI:10.1098/rsif.2013.0216 
  • Curtis N, Jones MEH, Evans SE, O'Higgins P, Fagan MJ. 2013. Cranial sutures work collectively to distribute strain throughout the reptile skul. Journal of Royal Society Interface 10: DOI:10.1098/rsif.2013.0442
  • Dong L, Roček Z, Wang Y, Jones MEH. 2013.  Anurans from the Lower Cretaceous Jehol Group of Western Liaoning, China. PLoS ONE 8(7): e69723. DOI:10.1371/journal.pone.0069723
  • Pearson M, Jones MEH. 2013. 7th World Congress of Herpetology (meeting report). Palaeontological Association Newsletter 83: 64–68. http://www.palass.org
  • Khannoon E, Evans SE. 2013. The embryonic development of the Egyptian cobra Naja h. haje (Squamata: Serpentes: Elapidae). Acta Zoologica DOI: 10.1111/azo.12043
  • Matsumoto R, Buffetaut E, Escuille F, Hervet S, Evans SE. 2013. New material of the choristodere  Lazarussuchus (Diapsida, Choristodera) from the Paleocene of France. Journal of Vertebrate Paleontology 33(2):319-339.  DOI: 10.1080/02724634.2012.716274
  • Benson RBJ, Mannion PD, Butler RJ, Upchurch P, Goswami A, Evans SE. 2013. Cretaceous tetrapod record sampling: implications for biogeography and the rise of extant clades. Palaeogeography, Palaeoclimatology, Palaeoecology 372: 88-107. DOI: 10.1016/j.palaeo.2012.10.028
  • Pearson MR, Benson RBJ, Upchurch P, Fröbischc J, Kammerer CF. 2013. Reconstructing the diversity of early terrestrial herbivorous tetrapods. Palaeogeography, Palaeoclimatology, Palaeoecology 372: 41-49. DOI: 10.1016/j.palaeo.2012.11.008
  • Jones MEH, Anderson CL, Hipsley CA, Müller J, Evans SE, Schoch R. 2013. Integration of molecules and  new fossils supports a Triassic origin for Lepidosauria (lizards, snakes, and  tuatara). BMC Evolutionary Biology 13:208 DOI:10.1186/1471-2148-13-208


  • Nesbitt SJ, Barrett PM, Werning S, Sidor CA, Charig AJ. 2012. The oldest dinosaur? A Middle Triassic dinosauriform from Tanzania. Biology Letters DOI: 10.1098/rsbl.2012.0949
  • Jones MEH, O'Higgins P, Fagan M, Evans SE, Curtis N. 2012. Shearing mechanics and the influence of a sflexible ymphysis during oral food processing in Sphenodon (Lepidosauria: Rhynchocephalia). The Anatomical Record 295: 1075-1091. DOI 10.1002/ar.22487
  • Evans SE, Wang Y. 2012. New material of the Early Cretaceous lizard Yabeinosaurus  from China. Cretaceous Research 34: 48–60.  doi:10.1016/j.cretres.2011.10.004
  • Mo JY, Xu X , Evans SE. 2012. A large predatory lizard (Platynota, Squamata) from the Late Cretaceous of South China. Journal of Systematic Palaeontology 10: 333–339. doi: 10.1080/14772019.2011.588254
  • Bolet A, Evans SE. 2012. A tiny lizard (Lepidosauria, Squamata) from the Lower Cretaceous of Spain. Palaeontology 55: 491–500. doi: 10.1111/j.1475-4983.2012.01145.x
  • Evans DC, Barrett PM, Seymour KL. 2012. Revised identification of a reported Iguanodon-grade ornithopod tooth from the Scollard Formation, Alberta, Canada Cretaceous Research 33: 11–14. DOI: 10.1016/j.cretres.2011.07.002
  • Bates KT, Maidment SCR, Allen V, Barrett PM. 2012. Computational modelling of locomotor muscle moment arms in the basal dinosaur Lesothosaurus diagnosticus: assessing convergence between birds and basal ornithischians. Journal of Anatomy 220: 212–232. DOI: 10.1111/j.1469-7580.2011.01469.x
  • Apesteguía S, Jones MEH. 2012. A Late Cretaceous "tuatara" (Lepidosauria: Sphenodontinae) from South America. Cretaceous Research 34: 154–160 doi:10.1016/j.cretres.2011.10.014
  • Bennett SP, Barrett PM, Collinson ME, Moore-Fay S, Davis PG, Palmer CP. 2012. A new specimen of Ichthyosaurus communis from Dorset, UK, and its bearing on the stratigraphical range of the species. Proceedings of the Geologists' Association 123: 146–154 . DOI: 10.1016/j.pgeola.2011.07.001
  • Butler RJ, Barrett PM, Gower DJ. 2012. Reassessment of the evidence for postcranial skeletal pneumaticity in Triassic archosaurs, and the early evolution of the avian respiratory system. PLoS ONE 7(3): e34094 . DOI: 10.1371/journal.pone.0034094
  • Smith AB, Barrett PM. 2012. Modelling the past: new generation approaches to understanding biological patterns in the fossil record: Introduction. Biology Letters 8: 112–114. DOI: 10.1098/rsbl.2011.1051
  • Smith AB, Barrett PM. (eds)  2012. Modelling the past: new generation approaches to understanding biological patterns in the fossil record. Biology Letters 8: 112–159. Special Feature
  • Mannion PD, Upchurch P, Mateus O, Barnes RN, Jones MEH. 2012. New information on the anatomy and systematic position of Dinheirosaurus lourinhanensis (Sauropoda: Diplodocoidea) from the Late Jurassic of Portugal, with a review of European diplodocoids. Journal of Systematic Palaeontology 10: 521-551. DOI: 10.1080/14772019.2011.595432
  • Meloro C, Jones MEH. 2012. Tooth and cranial disparity in the fossil relatives of Sphenodon (Rhynchocephalia) dispute the persistent ‘living fossil’ label. Journal of Evolutionary Biology 11: 2194–2209. DOI: 10.1111/j.1420-9101.2012.02595.x
  • Jones MEH, Werneburg I, Curtis N, Penrose R, O’Higgins P, Fagan MJ, Evans SE. 2012. The head and neck anatomy of sea turtles (Cryptodira: Chelonioidea) and skull shape in Testudines. PLoS ONE 7(11): e47852 DOI:10.1371/journal.pone.0047852
  • Evans SE, Jones MEH, Matsumoto R. 2012. A new lizard skull from the Purbeck Limestone Group (Lower Cretaceous) of England. Bulletin de la Société Géologique de France 183: 517–524. DOI: 10.2113/​gssgfbull.183.6.517
  • Benson RBJ, Mannion PD, Upchurch P, Goswami A, Evans SE. 2012. Cretaceous tetrapod fossil record sampling and faunal turnover: Implications for biogeography and the rise of modern clades. Palaeogeography, Palaeoclimatology, Palaeoecology DOI: 10.1016/j.palaeo.2012.10.028
  • Jones MEH, Cree A. 2012. Tuatara. Current Biology 22: 986–987 DOI: 10.1016/j.cub.2012.10.049


  • Wang Y, Evans SE. 2011. A gravid lizard from the Cretaceous of China and the early history of squamate viviparity. Naturwissenschaften DOI: 10.1007/s00114-011-0820-1
  • Barrett PM, Benson RBJ, Rich TH, Vickers-Rich P. 2011. First spinosaurid dinosaur from Australia and the cosmopolitanism of Cretaceous dinosaur faunas. Biology Letters DOI: 10.1098/rsbl.2011.0466
  • Evans SE, Wang Y. 2011. A new lizard (Reptilia: Squamata) with exquisite preservation of soft tissue from the Lower Cretaceous of Inner Mongolia, China. Journal of Systematic Palaeontology 8: 81-95. DOI: 10.1080/14772010903537773
  • Fortuny J, Bolet A, Sellés AG, Cartanyà J, Galobart À. 2011. New insights on Permian and Triassic vertebrates from north Eastern Iberian Peninsula. Journal of Iberian Geology 37(1): 65-86.
  • Kieser JA, He L-H, Dean MC, Jones MEH, Duncan WJ, Swain MV, Nelson NJ. 2011. Structure and compositional characteristics of caniniform dental enamel in the tuatara Sphenodon punctatus (Lepidosauria: Rhynchocephalia). New Zealand Dental Journal 107: 44-50.
  • Jones MEH, Curtis N, Fagan MJ, O’Higgins P, Evans SE. 2011. Hard tissue anatomy of the cranial joints in Sphenodon (Rhynchocephalia): sutures, kinesis, and skull mechanics. Palaeontologia Electronica 14, Issue 2; 17A:92p; palaeo-electronica.org/2011_2/251/index.html
  • Evans SE, Wang Y. 2011. New material of the Early Cretaceous lizard Yabeinosaurus  from China. Cretaceous Research 34: 48-60.  doi:10.1016/j.cretres.2011.10.004
  • Curtis N, Jones MEH, Shi J, O'Higgins P, Evans SE, Fagan MJ. 2011. Functional relationship between skull form and feeding mechanics in Sphenodon, and implications for diapsid skull development. PLoS ONE 6(12): e29804. doi:10.1371/journal.pone.0


  • Benson RBJ, Barrett PM, Rich TH, Vickers-Rich P. 2010 A southern tyrant reptile. Science 327: 1613.  DOI 10.1126/science.1187456
  • Matsumoto R, Evans SE. 2010. Choristoderes and the freshwater assemblages of Laurasia. Journal of Iberian Geology 36: 253-274. Weblink
  • Anquetin J. 2010. The anatomy of the basal turtle Eileanchelys waldmani from the Middle Jurassic of the Isle of Skye, Scotland. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 101: 67–96.  DOI: 10.1017/S1755691010009217
  • Müller J, Scheyer TM, Head JJ, Barrett PM, Werneburg I, Ericson PGP, Pol D, Sánchez-Villagra M R. 2010. Homeotic effects, somitogenesis and the evolution of vertebral numbers in recent and fossil amniotes. Proceedings of the National Academy of Sciences, PNAS 107, 2118­2123. DOI: 10.1073/pnas.0912622107
  • Evans SE, Wang Y. 2010. A new lizard (Reptilia: Squamata) with exquisite preservation of soft tissue from the Lower Cretaceous of Inner Mongolia, China. Journal of Systematic Palaeontology 8: 81-95. DOI: 10.1080/14772010903537773
  • Curtis N, Jones MEH, Evans SE, Shi J, O’Higgins P, Fagan MJ. 2010. Predicting muscle activation patterns from motion and anatomy: modelling the skull of Sphenodon. Royal Society Interface 7: 153–160 DOI: 10.1098/rsif.2009.0139
  • Bolet A, Evans SE. 2010. A new lizard from the Early Cretaceous of Catalonia (Spain), and the Mesozoic lizards of the Iberian Peninsula. Cretaceous Research 31: Pages 447-457. DOI: 10.1016/j.cretres.2010.06.002
  • Curtis N, Jones MEH, Lappin AK, Evans SE, O’Higgins P, Fagan MJ. 2010. Comparison between in vivo and theoretical bite performance: Using multi-body modelling to predict muscle and biteforces in a reptile skull. Journal of Biomechanics 43: 2804-2809. DOI: 10.1016/j.jbiomech.2010.05.037
  • Curtis N, Jones MEH, Evans SE, O’Higgins P, Fagan MJ. 2010. Feedback control from the jaw joints during biting: an investigation of the reptile Sphenodon using multibody modelling. Journal of Biomechanics 43: 3132-3137. DOI:10.1016/j.jbiomech.2010.08.001
  • Mo J-Y, Xu-X, Evans SE. 2010. The evolution of the lepidosaurian lower temporal bar: new perspectives from the Late Cretaceous of South China. Proceedings of the Royal Society B (Special Issue ‘Recent advances in Chinese palaeontology’) 277: 331-336. DOI: 10.1098/rspb.2009.0030
  • Evans SE, Jones MEH. 2010. The Origin, early history and diversification of lepidosauromorph reptiles. In Bandyopadhyay S. (ed.), New Aspects of Mesozoic Biodiversity, 27 Lecture Notes in Earth Sciences 132, 27-44. DOI 10.1007/978-3-642-10311-7_2
  • Müller J, Scheyer TM, Head JJ, Barrett PM, Werneburg I, Ericson PGP, Pol D, Sánchez-Villagra MR. 2010. Homeotic effects, somitogenesis and the evolution of vertebral numbers in recent and fossil amniotes. Proceedings of the National Academy of Sciences of America, PNAS  107: 2118–2123. DOI 10.1073/pnas.0912622107
  • Wang Y, Dong L, Evans SE. 2010. Jurassic-Cretaceous herpetofaunas from the Jehol associated strata in NE China: evolutionary and ecological implications. Bulletin of the Chinese Academy of Sciences 24: 76-79. Weblink
  • Archibald JD, Clemens WA, Padian K, Rowe T, MacLeod N, Barrett PM, Gale A, Holroyd P, Sues H-D, Arens NC, Horner JR, Wilson GP, Goodwin MB, Brichu CA, Lofgren DL, Hurlbert SH, Hartman JH, Eberth SA, Wignall PB, Currie PJ, Weil A, Prasad GVR, Dingus L, Courtillot V, Milner A, Milner A, Bajpai S, Ward DJ, Sahni A. 2010. Cretaceous extinctions: multiple causes. Science 328: 973. DOI: 10.1126/science.328.5981.973-a


  • Moazen M, Curtis N, O'Higgins P, Jones MEH, Evans SE, Fagan MJ, 2009. Assessment of the role of sutures in a lizard skull: a computer modelling study. Proceedings of the Royal Society B 276(1654): 39–46. DOI: 10.1098/rspb.2008.0863
  • Anquetin J, Barrett PM, Jones MEH, Moore-Fay S, Evans SE. 2009. A new stem turtle from the Middle Jurassic of Scotland: new insights into the evolution and palaeoecology of basal turtles. Proceedings of the Royal Society B 276: 879–886. DOI: 10.1098/rspb.2008.1429
  • Matsumoto R, Suzuki S, Tsogtbaatar K, Evans SE. 2009. New material of the enigmatic reptile Khurendukhosaurus (Diapsida: Choristodera) from Mongolia. Naturwissenschaften 96 (2): 233–242 http://www.springerlink.com/content/k05k3101vw24w832/
  • Walsh SA, Barrett PM, Milner AC, Manley G, Witmer LM. 2009. Inner ear anatomy is a proxy for deducing auditory capability and behaviour in reptiles and birds. Proceedings of the Royal Society B DOI: 10.1098/rspb.2008.1390
  • Jones MEH, Tennyson AJD, Worthy JP, Evans SE, Worthy TH. 2009. A sphenodontine (Rhynchocephalia) from the Miocene of New Zealand and palaeobiogeography of the tuatara (Sphenodon). Proceedings of the Royal Society B 276: 1385–1390 DOI: 10.1098/rspb.2008.1785
  • Evans SE, Wang Y. 2009. A long-limbed lizard from the Upper Jurassic/Lower Cretaceous of Daohugou, Ningcheng, Nei Mongol, China. Vertebrata PalAsiatica 47: 21–34.
  • Barrett PM, Butler RJ, Xiao-Lin W, Xing X. 2009. Cranial anatomy of the iguanodontoid ornithopod Jinzhousaurus yangi from the Lower Cretaceous Yixian Formation of China. Acta Palaeontologica Polonica 54: 35–48. http://www.app.pan.pl/article/item/app54-035.html
  • Zhang G, Wang Y, Jones MEH, Evans SE. 2009. A new Early Cretaceous salamander (Regalerpeton weichangensis gen. et sp. nov.) from the Huajiying Formation of northeastern China. Cretaceous Research 30: 51–558.DOI: 10.1016/j.cretres.2008.10.004
  • Moazen M, Curtis N, O’Higgins P, Evans SE, Fagan M. 2009. The function of the lower temporal bar in lepidosaurian evolution. Proceedings of the National Academy of Sciences. PNAS 20: 8273-8277. DOI: 10.1073/pnas.0813156106
  • Head JJ, Barrett PM, Rayfield EJ. 2009. Neurocranial osteology and systematic relationships of Varanus (Megalania) prisca Owen, 1859 (Squamata: Varanidae). Zoological Journal of the Linnean Society 155: 445–457. DOI: 10.1111/j.1096-3642.2008.00448.x
  • Jones MEH, Curtis N, O'Higgins P, Fagan M, Evans SE. 2009. Head and neck muscles associated with feeding in Sphenodon (Reptilia: Lepidsauria: Rhynchocephalia). Palaeontologia Electronica 12 (2, 7A): 1–56. Weblink
  • Manning PL, Margetts L, Johnson MR, Withers PJ, Sellers WI, Falkingham PL, Mummery PM, Barrett PM, Raymont DR. 2009. Biomechanics of dromaeosaurid dinosaur claws: application of X-Ray microtomography, nanoindentation, and finite element analysis. Anatomical Record 291:1397–1405. Weblink
  • Jones MEH, Lappin AK. 2009. Bite-force performance of the last rhynchocephalian (Lepidosauria: Sphenodon). Journal of the Royal Society of New Zealand 39(3): 71–83 Weblink
  • Barrett PM. 2009. The affinities of the enigmatic dinosaur Eshanosaurus deguchiianus from the Early Jurassic of Yunnan Province, People's Republic of China. Palaeontology 52: 681­688. DOI 10.1111/j.1475-4983.2009.00887
  • Barrett PM, McGowan AJ, Page V. 2009. Dinosaur diversity and the rock record. Proceedings of the Royal Society of London, Series B 276: 2667­2674. DOI: 10.1098/rspb.2009.0352
  • Butler RJ, Barrett PM, Gower DJ. 2009. Postcranial pneumaticity and bird-like air-sacs in the earliest pterosaurs. Biology Letters 5: 557­560. DOI: 10.1098/rsbl.2009.0139
  • Butler RJ, Barrett PM, Kenrick P, Penn MG. 2009. Diversity patterns amongst herbivorous dinosaurs and plants during the Cretaceous: implications for hypotheses of dinosaur/angiosperm co-evolution. Journal of Evolutionary Biology 22: 446­459. DOI 10.1111/j.1420-9101.2008.01680.x
  • Butler RJ, Barrett PM, Nowbath S, Upchurch P. In press.Estimating the effects of the rock record on pterosaur diversity patterns: implications for hypotheses of bird/pterosaur competitive replacement. Paleobiology 35: 432-446. Weblink
  • Rayfield EJ, Barrett PM, Milner AR. 2009. Utility and validity of Middle and Late Triassic 'Land Vertebrate Faunachrons'. Journal of Vertebrate Paleontology 29: 80­87. DOI: 10.1671/039.029.0132
  • Williams VS, Barrett PM, Purnell MA. 2009. Quantitative analysis of dental microwear in hadrosaurid dinosaurs, and the implications for hypotheses of jaw mechanics and feeding. Proceedings of the National Academy of Sciences, USA 106: 11194­11199. Weblink
  • Jones MEH. 2009. Dentary tooth shape in Sphenodon and its fossil relatives (Diapsida: Lepidosauria: Rhynchocephalia). In Koppe T, Meyer G, Alt KW, eds. Interdisciplinary Dental Morphology, Frontiers of Oral Biology (vol 13). Griefswald, Germany; Karger. 9-15.
  • Curtis N, Jones MEH, Evans SE, O’Higgins P, Fagan MJ. 2009. Visualising muscle anatomy using three-dimensional computer models - An example using the head and neck muscles of Sphenodon. Palaeontologia Electronica 12, Issue 3, 7T: 18p Weblink
  • Evans SE, Borsuk-Bialynicka M. 2009. The Early Triassic stem-frog Czatkobatrachus from Poland. Palaeontologica Polonica 65: 79–105. Weblink
  • Evans SE. 2009. An early kuehneosaurid reptile (Reptilia: Diapsida) from the Early Triassic of Poland. Palaeontologica Polonica 65: 145-178. Weblink
  • Evans SE, Borsuk-Bialynicka M. 2009. A small lepidosauromorph reptile from the Early Triassic of Poland. Palaeontologica Polonica 65: 179–202. Weblink
  • Borsuk-Bialynicka M, Evans SE. 2009. A long-necked archosauromorph from the Early Triassic of Poland. Palaeontologica Polonica 65: 203–234. Weblink
  • Borsuk-Bialynicka M, Evans SE. 2009. Cranial and mandibular osteology of the Early Triassic archosauriform Osmolskina czatkowicensis from Poland. Palaeontologica Polonica 65: 235–281. Weblink


  • Evans SE, Jones MEH, Krause DW. 2008. A giant frog with South American affinities from the Late Cretaceous of Madagascar. Proceedings of the National Academy of Sciences, PNAS 105: 2951–2956. DOI: 10.1073/pnas.0707599105
  • Whiteside DI, Marshall JEA. 2008. The age, fauna and palaeoenvironment of the Late Triassic fissure deposits of Tytherington, South Gloucestershire, UK. Geological Magazine 145:105–147. DOI: 10.1017/S0016756807003925
  • Scheyer TM, Anquetin J. 2008. Bone histology of the Middle Jurassic turtle shell remains from Kirtlington, Oxfordshire, England. Lethaia 41(1): 85–96. DOI: 10.1111/j.1502-3931.2007.00044.x
  • Evans SE, Manabe M. 2008. An early herbivorous lizard from the lower Cretaceous of Japan. Palaeontology. 51(2): 487–498. DOI: 10.1111/j.1475-4983.2008.00759.x
  • Sadleir RW, Barrett PM, Powell HP. 2008. The anatomy and systematics of Eustreptospondylus oxoniensis, a theropod dinosaur from the Middle Jurassic of Oxfordshire, England. Palaeontographical Society Monographs 160 (number 627): 1-82 + 20 pls.
  • Moazen M, Curtis N, Evans SE, O’Higgins P, Fagan M. 2008. Rigid-body analysis of a lizard skull: Modelling the skull of Uromastyx hardwickii. Journal of Biomechanics 41:1274–1280. DOI: 10.1016/j.jbiomech.2008.01.012 0
  • Jones MEH. 2008. Skull shape and feeding strategy in Sphenodon and other Rhynchocephalia (Diapsida: Lepidosauria). Journal of Morphology. 269: 945–-966. DOI: 10.1002/jmor.10634
  • Anquetin J, Claude J. 2008. A reassessment of the oldest British turtle: ‘Protochelys’ from the Middle Jurassic Stonesfield Slate of Stonesfield, Oxfordshire, UK. Geodiversitas 30: 331–344. Weblink
  • Buscalioni AD, Fregenal MA, Bravo A, Poyato-Ariza FJ, Sanchíz B, Báez AM, Cambra Moo O, Martín Closas C, Evans SE, Marugán Lobón J. 2008. The vertebrate assemblage of Buenache de la Sierra (Upper Barremian of Serrania de Cuenca, Spain) with insights into its taphonomy and palaeoecology. Cretaceous Research 29(4): 687–710. DOI:10.1016/j.cretres.2008.02.004
  • Moazen M, Curtis N, Evans SE, O'Higgins P, Fagan MJ. 2008. Combined finite element and multibody dynamics analysis of biting in a Uromastyx hardwickii lizard skull. Journal of Anatomy 213(5): 499–508. DOI: 10.1111/j.1469-7580.2008.00980.x
  • Evans SE. 2008. The skull of lizards and tuatara. In Biology of the Reptilia, Vol.20, Morphology H: the skull of Lepidosauria, Gans C, Gaunt A S, Adler K. (eds). Ithica, New York, Society for the study of Amphibians and Reptiles. pp1–344. Weblink to purchase