2021 | MRC Investigator

2017 | Associate Professor, MRC LMCB, UCL, London

2013 | Group Leader, MRC LMCB, UCL, London

2007-2013 | Senior Research Investigator, Cornell University, USA 

2004-2006 | HHMI Research Specialist, UCSD School of Medicine, USA

1998-2003 | Postdoctoral Fellow, UCSD School of Medicine, USA

1991-1998 | PhD, Washington University School of Medicine, USA

1986-1990 | BSc, with Honours, University of Nebraska, USA