2007 | Group Leader, MRC Laboratory for Cell Biology and  Department of Anatomy, UCL

2001-2007 | Royal Society URF at UCL and Group Leader at the UCL branch of the Ludwig Institute for Cancer Research

1997-2001 | Postdoctoral Fellow, Harvard Medical School, USA

1993 | PhD, UCL, UK

1989 | Biochemistry, 1st Class Hons, St Catherine's College, Oxford, UK