2002- | Emeritus Professor, MRC Laboratory for Molecular Cell Biology , UCL

1993-2002 | Scientist, MRC Laboratory for Molecular Cell Biology, University College London

1979-2002 | Professor of Biology, Biology Department, University College London

1971-2002|  MRC Developmental Neurobiology Programme, Biology Department, University College London

1969-1971 | Postdoctoral Fellow in Immunology, National Institute for Medical Research, Mill Hill, London

1965-1968 | Resident in Neurology, Massachusetts General Hospital, Boston

1963-1965 | Intern and Assistant Resident in Medicine, Royal Victoria Hospital, Montreal

2016 | Elected Fellow of AAAS

2010 | DART/NYU Biotechnology Achievement Award in Basic Science

2007 | Honorary Degree, Vrije Universiteit Brussel

2006 | The Biochemical Society Award

2005 | Honorary Degree, McGill University

2005 | Honorary Fellow, University College London

2004 | Kuffler Lectures, UCSD

2003 | Foreign Associate of National Academy of Sciences of the USA

2002 | Hamdan Prize for Apoptosis in Disease and Health

2002 | United Kingdom-Canada Rutherford Lectures

2001 | Dunham Lectures, Harvard University

1999 | Foreign Honorary Member of the American Academy of Arts and Sciences

1998 | British Academy of Medical Sciences

1997–2001 | Chairman, UK Life Sciences Committee

1991–1995 | President, British Society of Cell Biology

1989 | Honorary Member of American Neurological Society

1989 | Feldberg Prize

1988 | Academia Europaea

1985 | Fellow of the Royal Society

1974 | Member European Molecular Biology Organization