1992 | Group Leader, MRC Laboratory for Molecular Cell Biology, UCL, London, UK

1985-1991 | Group Leader, Institute of Cancer Research, London, UK

1981-1985 | Postdoctoral Fellow, Yale University Medical School, USA

1979-1981 | Postdoctoral Fellow, EMBL, Heidelberg, Germany

1975-1978 | PhD, Zoology UCL, UK

1972-1975 | BSc, Zoology UCL, UK