Heliconiina checklist with links to maps. Modified from Lamas 2004, synonyms excluded, with some revised placements and additions. J. Mallet, July 2011

Checklist © Gerardo Lamas, 2004. Please cite: Lamas, G., C. Callaghan, M. M. Casagrande, O. Mielke, T. Pyrcz, R. Robbins and A. Viloria. 2004. Hesperioidea -- Papilionoidea. In: Heppner, J.B. (ed.) Atlas of Neotropical Lepidoptera. Checklist: part 4A. Association for Tropical Lepidoptera/Scientific Publishers, Gainesville, Florida. Updated REPRINT

Distribution data © Neil Rosser, 2012. Full data is available at http://www.ucl.ac.uk/taxome/neil_rosser/
Please cite: Rosser, N., Phillimore, A.B., Huertas, B., Willmott, K.R., & Mallet, J. (2012). Testing historical explanations for gradients in species richness in heliconiine butterflies of tropical America. Biological Journal of the Linnean Society 105: 479-497. REPRINT

Genus
Species Subspecies
Author
Date
 Original genus Type Locality
Agraulis



J.B.A.D. Boisduval & J.E. Le Conte
1835



vanillae

C. Linnaeus
1758
 Papilio



vanillae
C. Linnaeus
1758
 Papilio America



forbesi
C.D. Michener
1942
Peru



galapagensis
W.J. Holland
1890
Galapagos



incarnata
N.D. Riley
1926
 Dione Mexico (Dur)



insularis
C.J. Maynard
1889
Bahamas



lucina
C. Felder & R. Felder
1862
"Brazil (Am)"



maculosa
H.F.E.J. Stichel
1908
 Dione Brazil (ES); Paraguay



nigrior
C.D. Michener
1942
USA (Fl)


[n. sp.]

G. Lamas
MS
Peru
Dione



J. Hübner
1819



glycera

C. Felder & R. Felder
1861
 Agraulis Venezuela; Colombia


juno

P. Cramer
1779
 Papilio



juno
P. Cramer
1779
 Papilio Surinam



andicola
H.W. Bates
1864
 Agraulis Ecuador



huascuma
T. Reakirt
1866
 Agraulis Mexico



miraculosa
M.E.R. Hering
1926
Peru



suffumata
K.S. Brown & O. Mielke
1972
Paraguay


moneta

J. Hübner
1825




moneta
J. Hübner
1825
?



butleri
H.F.E.J. Stichel
1908
Colombia



poeyii
A.G. Butler
1873
Mexico [(Oax)]
Podotricha



C.D. Michener
1942



judith

F.E. Guérin-Méneville
1844
 Cethosia



judith
F.E. Guérin-Méneville
1844
 Cethosia Colombia



caucana
N.D. Riley
1926
 Colaenis Colombia



mellosa
H.F.E.J. Stichel
1906
 Colaenis Ecuador



straminea
N.D. Riley
1926
 Colaenis Ecuador


telesiphe

W.C. Hewitson
1867
 Colaenis



telesiphe
W.C. Hewitson
1867
 Colaenis Ecuador



tithraustes
O. Salvin
1871
 Colaenis Ecuador
Dryadula



C.D. Michener
1942



phaetusa

C. Linnaeus
1758
 Papilio "Indiis"
Dryas



J. Hübner
1807



iulia

J.C. Fabricius
1775
 Papilio



iulia
J.C. Fabricius
1775
 Papilio America



alcionea
P. Cramer
1779
 Papilio Surinam



carteri
N.D. Riley
1926
 Colaenis Bahamas



delila
J.C. Fabricius
1775
 Papilio America



dominicana
A. Hall
1917
 Colaenis Dominica



framptoni
N.D. Riley
1926
 Colaenis St. Vincent



fucatus
P. Boddaert
1783
 Papilio Dominican Rep.



lucia
N.D. Riley
1926
 Colaenis St. Lucia



martinica
P. Enrico & R. Pinchon
1969
Martinique



moderata
N.D. Riley
1926
 Colaenis Honduras



nudeola
M. Bates
1934
 Colaenis Cuba



warneri
A. Hall
1936
 Colaenis St. Kitts



zoe
L.D. Miller & S.R. Steinhauser
1992
Cayman Islands
Philaethria



G.J. Billberg
1820


Metamandana


H.F.E.J. Stichel
1908



andrei

C. Brévignon
2002
French Guiana



andrei







orinocensis
J.M. Constantino & J.A. Salazar
2010
Venezuela


browni

J.M. Constantino & J.A. Salazar
2010
Venezuela


constantinoi

J.A. Salazar
1991
Colombia


diatonica

H. Fruhstorfer
1912
 Metamandana Honduras


dido

C. Linnaeus
1763
 Papilio



dido
C. Linnaeus
1763
 Papilio S. Amer.



chocoensis
L.M. Constantino
1999
Colombia



panamensis
J.M. Constantino & J.A. Salazar
2010
Panama


neildi

J.M. Constantino & J.A. Salazar
2010




neildi
J.M. Constantino & J.A. Salazar
2010
Venezuela



tachiraensis
J.M. Constantino & J.A. Salazar
2010
Venezuela



winhardi
J.M. Constantino & J.A. Salazar
2010
Colombia


ostara

J.K.M. Röber
1906
 Metamorpha



ostara
J.K.M. Röber
1906
 Metamorpha Colombia



araguensis
J.M. Constantino & J.A. Salazar
2010
Venezuela



meridensis
J.M. Constantino & J.A. Salazar
2010
Venezuela


pygmalion

H. Fruhstorfer
1912
 Metamandana



pygmalion
H. Fruhstorfer
1912
 Metamandana Brazil (Pa)



metaensis
J.M. Constantino & J.A. Salazar
2010
Colombia


romeroi

J.M. Constantino & J.A. Salazar
2010
Venezuela


wernickei

J.K.M. Röber
1906
 Metamorpha Brazil (RS/SC)
Laparus



G.J. Billberg
1820


Crenis


J. Hübner
1821



doris

C. Linnaeus
1771
 Papilio



doris
C. Linnaeus
1771
 Papilio Surinam



dives
C. Oberthür
1920
 Heliconia Colombia



obscurus
G. Weymer
1891
 Heliconius Colombia



viridis
O. Staudinger
1885
 Heliconius Panama



[n. ssp.]
W.M. Neukirchen
MS
Trinidad
Neruda



J.R.G. Turner
1976



aoede

J. Hübner
1813
 Nerëis



aoede
J. Hübner
1813
 Nerëis [Brazil]



aliciae
W.M. Neukirchen
2000
Ecuador



astydamia
W.F. Erichson
1849
 Heliconia Guyana



auca
W.M. Neukirchen
1997
Ecuador



ayacuchensis
W.M. Neukirchen
1992
 Heliconius (Neruda) Venezuela



bartletti
H. Druce
1876
 Heliconius Peru



centurius
W.M. Neukirchen
1994
 Heliconius (Neruda) French Guiana



cupidineus
H.F.E.J. Stichel
1906
 Heliconius Peru



emmelina
C. Oberthür
1902
 Heliconia Guyana



eurycleia
K.S. Brown
1973
 Heliconius Brazil (MT)



faleria
H. Fruhstorfer
1910
 Heliconius Brazil (MT)



lucretius
G. Weymer
1891
 Heliconius Brazil (Am)



manu
G. Lamas
1976
 Heliconius Peru



philipi
K.S. Brown
1976
 Heliconius Bolivia



[n. ssp.]
K.S. Brown
MS
Brazil


godmani

O. Staudinger
1882
 Heliconius Colombia


metharme

W.F. Erichson
1849
 Heliconia



metharme
W.F. Erichson
1849
 Heliconia Guyana



makiritare
K.S. Brown & F. Fernández
1985
Venezuela



perseis
H.F.E.J. Stichel
1923
 Heliconius Colombia


metis

G.R.P. Moreira & C.G.C. Mielke
2010
Brazil
Eueides



J. Hübner
1816



aliphera

J.B. Godart
1819
 Cethosia



aliphera
J.B. Godart
1819
 Cethosia Brazil



cyllenella
A. Seitz
1912
?



gracilis
H.F.E.J. Stichel
1903
Honduras; Costa Rica


emsleyi

K.S. Brown
1976




emsleyi
K.S. Brown
1976
Colombia



[n. ssp.]
K.S. Brown
MS
Ecuador


heliconioides

C. Felder & R. Felder
1861




heliconioides
C. Felder & R. Felder
1861
"Ecuador"



eanes
W.C. Hewitson
1861
Peru



koenigi
H. Holzinger & R. Holzinger
1993
Peru


isabella

C. Stoll
1781
 Papilio



isabella
C. Stoll
1781
 Papilio Surinam



arquata
H.F.E.J. Stichel
1903
Colombia



cleobaea
C. Geyer
1832
Cuba



dianasa
J. Hübner
1806
 Nerëis Brazil (Ba)



dissoluta
H.F.E.J. Stichel
1903
Peru



dynastes
C. Felder & R. Felder
1861
Venezuela; Colombia



ecuadorensis
E. Strand
1912
Ecuador



eva
J.C. Fabricius
1793
 Papilio "Surinam"



hippolinus
A.G. Butler
1873
Peru



huebneri
E. Ménétriés
1857
"Brazil"



melphis
J.B. Godart
1819
 Heliconia Antilles



nigricornis
R.G. Maza
1982
Mexico (Gue)



[n. ssp.]
K.S. Brown
MS
Brazil (Ce)


lampeto

H.W. Bates
1862




lampeto
H.W. Bates
1862
Brazil (Am)



acacetes
W.C. Hewitson
1869
Ecuador



apicalis
J.K.M. Röber
1927
Ecuador



carbo
A. Seitz
1913
Ecuador



concisa
G. Lamas
1985
Peru



nigrofulva
W.J. Kaye
1906
Guyana



[n. ssp.]
K.S. Brown
MS
French Guiana



[n. ssp.]
K.S. Brown
MS
Brazil (Am)


libitina

O. Staudinger
1885




libitina
O. Staudinger
1885
French Guiana



malleti
G. Lamas
1998
Peru


lineata

O. Salvin & F.D. Godman
1868
Guatemala


lybia

J.C. Fabricius
1775
 Papilio



lybia
J.C. Fabricius
1775
 Papilio "India"



lybioides
O. Staudinger
1876
Panama



olympia
J.C. Fabricius
1793
 Papilio America



orinocensis
K.S. Brown & F. Fernández
1985
Venezuela



otelloi
K.S. Brown & F. Fernández
1985
Venezuela



salcedoi
K.S. Brown & F. Fernández
1985
Venezuela


pavana

E. Ménétriés
1857
Brazil


procula

E. Doubleday
1847




procula
E. Doubleday
1847
Venezuela



asidia
W. Schaus
1920
Guatemala



browni
H. Holzinger & R. Holzinger
1974
Venezuela



edias
W.C. Hewitson
1861
Colombia



eurysaces
W.C. Hewitson
1864
Ecuador



kuenowii
H. Dewitz
1877
Colombia



vulgiformis
A.G. Butler & H. Druce
1872
Costa Rica


tales

P. Cramer
1775
 Papilio



tales
P. Cramer
1775
 Papilio Surinam



barcellinus
J. Zikán
1937
Brazil (Am)



calathus
H.F.E.J. Stichel
1909
Ecuador



cognata
G. Weymer
1890
Colombia



franciscus
K.S. Brown & H. Holzinger
1973
Venezuela



michaeli
J. Zikán
1937
Peru



pseudeanes
E. Boullet & F.L. Le Cerf
1910
Venezuela



pythagoras
W.F. Kirby
1900
Brazil (Pa)



surdus
H.F.E.J. Stichel
1903
Brazil (Pa); Guyana



tabernula
G. Lamas
1985
Peru



xenophanes
C. Felder & R. Felder
1865
Colombia



[n. ssp.]
K.S. Brown
MS
French Guiana



[n. ssp.]
K.S. Brown
MS
Brazil (Pa)


vibilia

J.B. Godart
1819
 Cethosia



vibilia
J.B. Godart
1819
 Cethosia Brazil



mereaui
J. Hübner
1823
 Colaenis "Brazil"



unifasciatus
A.G. Butler
1873
Brazil (Am)



vialis
H.F.E.J. Stichel
1903
Costa Rica



vicinalis
H.F.E.J. Stichel
1903
Ecuador



[n. ssp.]
K.S. Brown
MS
Colombia
Heliconius



K. Kluk
1780


1 antiochus

C. Linnaeus
1767
 Papilio



antiochus
C. Linnaeus
1767
 Papilio "Indiis"



aranea
J.C. Fabricius
1793
 Papilio ?



araneides
O. Staudinger
1897
Venezuela



salvinii
H. Dewitz
1877
[Venezuela]



[n. ssp.]
G. Lamas
MS
Peru

2 astraea

O. Staudinger
1897




astraea
O. Staudinger
1897
Brazil (Am)



rondonia
K.S. Brown
1973
Brazil (Rd)

3 atthis

E. Doubleday
1847
 Heliconia Ecuador

4 besckei

E. Ménétriés
1857
 Heliconia Brazil (RJ/MG)

5 burneyi

J. Hübner
1831
 Migonitis



burneyi
J. Hübner
1831
 Migonitis Brazil



ada
H. Neustetter
1925
Brazil (Rd)



anjae
W.M. Neukirchen
1995
Brazil (Am)



boliviensis
W.M. Neukirchen
1995
Bolivia



catharinae
O. Staudinger
1885
[Brazil (Pa)]



huebneri
O. Staudinger
1897
[Peru]



jamesi
W.M. Neukirchen
1995
Peru



koenigi
W.M. Neukirchen
1995
Peru



lindigii
C. Felder & R. Felder
1865
Colombia



skinneri
K.S. Brown & F. Fernández
1985
Venezuela

6 charithonia

C. Linnaeus
1767
 Papilio



charithonia
C. Linnaeus
1767
 Papilio America



antiquus
G. Lamas
1988
St. Kitts; Antigua



bassleri
W.P. Comstock & F.M. Brown
1950
Colombia



churchi
W.P. Comstock & F.M. Brown
1950
Haiti



ramsdeni
W.P. Comstock & F.M. Brown
1950
Cuba



simulator
J.K.M. Röber
1921
Jamaica



tuckeri
W.P. Comstock & F.M. Brown
1950
USA (Fl)



vazquezae
W.P. Comstock & F.M. Brown
1950
Mexico (Cam)

7 chestertonii

W.C. Hewitson
1872
 Heliconia Colombia

8 clysonymus

P.A. Latreille
1817




clysonymus
P.A. Latreille
1817
[Colombia]



hygiana
W.C. Hewitson
1867
 Heliconia Ecuador



montanus
O. Salvin
1871
Costa Rica



tabaconas
K.S. Brown
1976
Peru

9 congener

G. Weymer
1890




congener
G. Weymer
1890
Ecuador



aquilionaris
K.S. Brown
1976
Colombia



ocannensis
H.F.E.J. Stichel
1905
Colombia

10 cydno

E. Doubleday
1847
 Heliconia



cydno
E. Doubleday
1847
 Heliconia Colombia



alithea
W.C. Hewitson
1869
 Heliconia Ecuador



barinasensis
J.H. Masters
1973
Venezuela



chioneus
H.W. Bates
1864
Panama



cordula
H. Neustetter
1913
Venezuela



cydnides
O. Staudinger
1885
Colombia



gadouae
K.S. Brown & F. Fernández
1985
Venezuela



galanthus
H.W. Bates
1864
Guatemala



hermogenes
W.C. Hewitson
1858
 Heliconia [Colombia]



lisethae
W.M. Neukirchen
1995
Colombia



pachinus [species] O. Salvin
1871
Panama



wanningeri
W.M. Neukirchen
1991
Colombia



weymeri
O. Staudinger
1897
Colombia



zelinde
A.G. Butler
1869
W coast of America

11 demeter

O. Staudinger
1897




demeter
O. Staudinger
1897
Peru



angeli
W.M. Neukirchen
1997
Peru



beebei
J.R.G. Turner
1966
Guyana



bouqueti
E. Nöldner
1901
French Guiana



eratosignis
J.J. Joicey & G. Talbot
1925
 Eueides Brazil (MT)



karinae
W.M. Neukirchen
1990
Brazil (Pa)



neildi
W.M. Neukirchen
1997
Ecuador



tambopata
G. Lamas
1985
Peru



terrasanta
K.S. Brown & W.W. Benson
1975
Brazil (Pa)



titan
W.M. Neukirchen
1995
Brazil (Am)



turneri
K.S. Brown & W.W. Benson
1975
Brazil (Am)



ucayalensis
H. Holzinger & R. Holzinger
1975
Peru



ulysses
K.S. Brown & W.W. Benson
1975
Bolivia



zikani
K.S. Brown & W.W. Benson
1975
Brazil (Am)



[n. ssp.]
G. Lamas
MS
Peru

12 egeria

P. Cramer
1775
 Papilio



egeria
P. Cramer
1775
 Papilio Surinam



christiani
W.M. Neukirchen
1997
Brazil (Pa)



egerides
O. Staudinger
1897
Brazil (Am)



homogena
F. Bryk
1953
Brazil (Am)



hyas
G. Weymer
1883
[Brazil (Am)]



keithbrowni
W.M. Neukirchen
1992
Venezuela



mariasibyllae
W.M. Neukirchen
1991
Brazil (Pa)

13 eleuchia

W.C. Hewitson
1854
 Heliconia



eleuchia
W.C. Hewitson
1854
 Heliconia Colombia



eleusinus
O. Staudinger
1885
Colombia



primularis
A.G. Butler
1869
Ecuador

14 elevatus

E. Nöldner
1901




elevatus
E. Nöldner
1901
"Amazonas"



bari
C. Oberthür
1902
 Heliconia French Guiana



lapis
G. Lamas
1976
Peru



perchlora
J.J. Joicey & W.J. Kaye
1917
Bolivia



pseudocupidineus
H. Neustetter
1931
Peru



roraima
J.R.G. Turner
1966
Guyana



schmassmanni
J.J. Joicey & G. Talbot
1925
Brazil (MT)



sonjae
W.M. Neukirchen
1997
Brazil (Pa)



taracuanus
F. Bryk
1953
Brazil (Am)



tumatumari
W.J. Kaye
1906
Guyana



willmotti
W.M. Neukirchen
1997
Ecuador



zoelleri
W.M. Neukirchen
1990
Venezuela

15 erato

C. Linnaeus
1758
 Papilio



erato
C. Linnaeus
1758
 Papilio Surinam



adana
J.R.G. Turner
1967
[Trinidad]



amalfreda
H.H. Riffarth
1901
Brazil (Pa)



amazona
O. Staudinger
1897
Brazil (Pa)



amphitrite
H.H. Riffarth
1901
Peru



chestertonii [species] W.C. Hewitson
1872
 Heliconia Colombia



colombina
O. Staudinger
1897
Colombia



cruentus
G. Lamas
1998
Mexico (Chia)



cyrbia
J.B. Godart
1819
 Heliconia America



demophoon
E. Ménétriés
1855
Nicaragua



dignus
H.F.E.J. Stichel
1923
Colombia



emma
H.H. Riffarth
1901
Peru; "Ecuador"



estrella
H.W. Bates
1862
Brazil (Pa)



etylus
O. Salvin
1871
Ecuador



favorinus
C.H. Hopffer
1874
"Bolivia"



guarica
T. Reakirt
1868
Colombia



hydara
W.C. Hewitson
1867
 Heliconia Colombia



lativitta
A.G. Butler
1877
Brazil (Am); Ecuador



lichyi
K.S. Brown & F. Fernández
1985
Venezuela



luscombei
G. Lamas
1976
Peru



magnifica
H.H. Riffarth
1900
Guyana



microclea
W.J. Kaye
1907
Peru



notabilis
O. Salvin & F.D. Godman
1868
Ecuador



petiverana
E. Doubleday
1847
 Heliconia Mexico



phyllis
J.C. Fabricius
1775
 Papilio Brazil



reductimacula
F. Bryk
1953
Brazil (Am)



tobagoensis
M. Barcant
1982
Tobago



venus
O. Staudinger
1882
Colombia



venustus
O. Salvin
1871
Bolivia

16 ethilla

J.B. Godart
1819
 Heliconia



ethilla
J.B. Godart
1819
 Heliconia "Antilles"



adela
H. Neustetter
1912
Peru



aerotome
C. Felder & R. Felder
1862
"Brazil (Am)"



cephallenia
C. Felder & R. Felder
1865
Surinam



chapadensis
K.S. Brown
1973
Brazil (MT)



claudia
F.D. Godman & O. Salvin
1881
Panama



eucoma
J. Hübner
1831
 Eueides Brazil



flavofasciatus
G. Weymer
1894
Brazil (Pa)



flavomaculatus
G. Weymer
1894
Brazil (Pe)



hyalina
H. Neustetter
1928
Brazil (Rr)



jaruensis
K.S. Brown
1976
Brazil (Rd)



latona
W.M. Neukirchen
1998
Colombia



mentor
G. Weymer
1883
Colombia



metalilis
A.G. Butler
1873
Venezuela



michaelianius
G. Lamas
1988
Peru



narcaea
J.B. Godart
1819
 Heliconia "Antilles"



nebulosa
W.J. Kaye
1916
Peru



neukircheni
G. Lamas
1998
Peru



numismaticus
G. Weymer
1894
Brazil (Pa)



penthesilea
W.M. Neukirchen
1994
Brazil (Am)



polychrous
C. Felder & R. Felder
1865
Brazil ([SP])



semiflavidus
G. Weymer
1894
Colombia



thielei
H.H. Riffarth
1900
French Guiana



tyndarus
G. Weymer
1897
Bolivia



yuruani
K.S. Brown & F. Fernández
1985
Venezuela










17 hecale

J.C. Fabricius
1776
 Papilio



hecale
J.C. Fabricius
1776
 Papilio Surinam



anderida
W.C. Hewitson
1853
 Heliconia [Venezuela]



annetta
H.H. Riffarth
1900
Colombia



australis
K.S. Brown
1976
Ecuador



barcanti
K.S. Brown
1976
Venezuela



clearei
A. Hall
1930
Guyana



discomaculatus
G. Weymer
1891
Honduras



ennius
G. Weymer
1891
Brazil (Am)



felix
G. Weymer
1894
Peru



fornarina
W.C. Hewitson
1854
 Heliconia ["S. Amer."]



holcophorus
O. Staudinger
1897
Colombia



humboldti
H. Neustetter
1928
Peru



ithaca
C. Felder & R. Felder
1862
Colombia



latus
H.H. Riffarth
1900
Brazil (Pa)



melicerta
H.W. Bates
1866
Panama; Colombia



metellus
G. Weymer
1894
Brazil (Pa)



naxos
W.M. Neukirchen
1998
Brazil (Pa)



nigrofasciatus
G. Weymer
1894
Brazil (Am)



novatus
H.W. Bates
1867
Brazil (Pa/Ma)



paraensis
H.H. Riffarth
1900
Brazil (Pa)



paulus
W.M. Neukirchen
1998
Brazil (Pa)



quitalena
W.C. Hewitson
1853
 Heliconia Ecuador



rosalesi
K.S. Brown & F. Fernández
1976
Venezuela



shanki
G. Lamas & K.S. Brown
1976
Peru



sisyphus
O. Salvin
1871
Peru



sulphureus
G. Weymer
1894
Brazil (Am)



vetustus
A.G. Butler
1873
Guyana



zeus
W.M. Neukirchen
1995
Bolivia



zuleika
W.C. Hewitson
1854
 Heliconia Nicaragua

18 hecalesia

W.C. Hewitson
1854
 Heliconia



hecalesia
W.C. Hewitson
1854
 Heliconia Colombia



eximius
H.F.E.J. Stichel
1923
Colombia



formosus
H.W. Bates
1866
Panama



gynaesia
W.C. Hewitson
1875
 Heliconia ?



longarena
W.C. Hewitson
1875
 Heliconia Colombia



octavia
H.W. Bates
1866
Guatemala



romeroi
K.S. Brown & F. Fernández
1985
Venezuela

19 hecuba

W.C. Hewitson
1858
 Heliconia



hecuba
W.C. Hewitson
1858
 Heliconia Colombia



bonplandi
W.M. Neukirchen
1991
Ecuador



cassandra
C. Felder & R. Felder
1862
Colombia



choarina
W.C. Hewitson
1872
 Heliconia Ecuador



creusa
H. Holzinger & R. Holzinger
1989
Colombia



crispus
O. Staudinger
1885
Colombia



flava
K.S. Brown
1979
Ecuador



lamasi
W.M. Neukirchen
1991
Ecuador



salazari
W.M. Neukirchen
1993
Colombia



tolima
A.H. Fassl
1912
Colombia



walteri
J.A. Salazar
1998
Colombia

20 hermathena

W.C. Hewitson
1854
 Heliconia



hermathena
W.C. Hewitson
1854
 Heliconia [Brazil]



duckei
K.S. Brown & W.W. Benson
1977
Brazil (Pa)



renatae
K.S. Brown & F. Fernández
1985
Venezuela



sabinae
W.M. Neukirchen
1992
Brazil (Am)



sheppardi
K.S. Brown & W.W. Benson
1977
Brazil (Am)



vereatta
H.F.E.J. Stichel
1912
Brazil (Pa)

21 heurippa

W.C. Hewitson
1854
 Heliconia



heurippa
W.C. Hewitson
1854
 Heliconia Colombia



[n. ssp.]
Linares et al.
MS
Colombia

22 hewitsoni

W.C. Hewitson
1875
 Heliconia Panama

23 hierax

W.C. Hewitson
1869
 Heliconia



hierax
W.C. Hewitson
1869
 Heliconia Ecuador



leveri
S. Attal
1999
Ecuador

24 himera

W.C. Hewitson
1867
 Heliconia Ecuador

25 hortense

F.E. Guérin-Méneville
1844
 Heliconia Mexico

26 ismenius

P.A. Latreille
1817




ismenius
P.A. Latreille
1817
[S. Amer.]



boulleti
H. Neustetter
1928
"French Guiana"



clarescens
A.G. Butler
1875
Panama



fasciatus
F.D. Godman & O. Salvin
1877
Panama



metaphorus
G. Weymer
1883
Ecuador



occidentalis
H. Neustetter
1928
Colombia



telchinia
E. Doubleday
1847
 Heliconia [Honduras]



tilletti
K.S. Brown & F. Fernández
1976
Venezuela

27 lalitae

C. Brévignon
1996
French Guiana

28 leucadia

H.W. Bates
1862




leucadia
H.W. Bates
1862
Brazil (Am)



andromeda
W.M. Neukirchen
1996
Ecuador



birgitae
W.M. Neukirchen
1996
Peru



pseudorhea
O. Staudinger
1897
Brazil (Am)

29 luciana

R. Lichy
1960




luciana
R. Lichy
1970
Venezuela



watunna
R. Lichy
1960
Venezuela

30 melpomene

C. Linnaeus
1758
 Papilio



melpomene
C. Linnaeus
1758
 Papilio America



aglaope
C. Felder & R. Felder
1862
"Brazil (Am)"



amandus
H. Grose-Smith & W.F. Kirby
1892
Bolivia



amaryllis
C. Felder & R. Felder
1862
"Brazil (Am)"



anduzei
K.S. Brown & F. Fernández
1985
Venezuela



bellula
J.R.G. Turner
1971
Colombia



burchelli
E.B. Poulton
1910
Brazil (Go)



cythera
W.C. Hewitson
1869
 Heliconia Ecuador



ecuadorensis
M.G. Emsley
1964
Ecuador



euryades
H.H. Riffarth
1900
Peru



flagrans
H.F.E.J. Stichel
1919
Trinidad



intersectus
H. Neustetter
1928
Brazil ([Pa])



madeira
N.D. Riley
1919
Brazil (Am)



malleti
G. Lamas
1988
Ecuador



meriana
J.R.G. Turner
1967
Surinam



michellae
W.M. Neukirchen
1997
Peru



nanna
H.F.E.J. Stichel
1899
Brazil (ES/MG)



penelope
O. Staudinger
1894
Bolivia



plesseni
H.H. Riffarth
1907
Ecuador



pyrforus
W.J. Kaye
1907
Guyana



rosina
J.B.A.D. Boisduval
1870
 Heliconia Costa Rica; "Mexico"



schunkei
G. Lamas
1976
Peru



tessa
M. Barcant
1982
Tobago



thelxiope
J. Hübner
1806
 Nerëis [Brazil]



thelxiopeia
O. Staudinger
1897
French Guiana



vicina
E. Ménétriés
1857
Brazil



vulcanus
A.G. Butler
1865
"Guyana"; Panama



xenoclea
W.C. Hewitson
1853
 Heliconia ?



[n. ssp.]
W.M. Neukirchen
MS
Colombia

31 nattereri

C. Felder & R. Felder
1865
Brazil ([Ba])

32 numata

P. Cramer
1780
 Papilio



numata
P. Cramer
1780
 Papilio Surinam



arcuella
H. Druce
1874
Peru



aristiona
W.C. Hewitson
1853
 Heliconia "Colombia"



aulicus
G. Weymer
1883
[Venezuela]



aurora
H.W. Bates
1862
Brazil (Am)



bicoloratus
A.G. Butler
1873
Peru



ethra
J. Hübner
1831
 Eueides Brazil



euphone
C. Felder & R. Felder
1862
Colombia



geminatus
G. Weymer
1894
Brazil (Am)



holzingeri
F. Fernández & K.S. Brown
1976
Venezuela



ignotus
J.J. Joicey & W.J. Kaye
1917
Peru



illustris
G. Weymer
1894
Peru



jiparanaensis
H. Neustetter
1931
Brazil (Rd)



lenaeus
G. Weymer
1891
"Colombia"



lyrcaeus
G. Weymer
1891
Peru?



mavors
G. Weymer
1894
[Brazil (Am)]



messene
C. Felder & R. Felder
1862
Colombia



mirus
G. Weymer
1894
Bolivia



nubifer
A.G. Butler
1875
Brazil (Am)



peeblesi
J.J. Joicey & G. Talbot
1925
Venezuela



pratti
J.J. Joicey & W.J. Kaye
1917
Peru



robigus
G. Weymer
1875
"Venezuela"



silvana
C. Stoll
1781
 Papilio Surinam



superioris
A.G. Butler
1875
Brazil (Am/Pa)



talboti
J.J. Joicey & W.J. Kaye
1917
Peru



tarapotensis
H.H. Riffarth
1901
Peru



zobrysi
H. Fruhstorfer
1910
Brazil (MT)

33 pachinus

O. Salvin
1871
Panama

34 pardalinus

H.W. Bates
1862




pardalinus
H.W. Bates
1862
Brazil (Am)



ariadne
W.M. Neukirchen
1995
Bolivia



butleri
K.S. Brown
1976
Peru



dilatus
G. Weymer
1894
Peru



julia
W.M. Neukirchen
2000
Ecuador



lucescens
G. Weymer
1894
Brazil (Pa)



maeon
G. Weymer
1891
?



orteguaza
K.S. Brown
1976
Colombia



radiosus
A.G. Butler
1873
Brazil (Pa)



sergestus
G. Weymer
1894
Peru



tithoreides
O. Staudinger
1900
Peru

35 peruvianus

C. Felder & R. Felder
1859
 Heliconia Peru

36 ricini

C. Linnaeus
1758
 Papilio



ricini
C. Linnaeus
1758
 Papilio America



insulanus
H.F.E.J. Stichel
1909
 Eueides Trinidad

37 sapho

D. Drury
1782
 Papilio



sapho
D. Drury
1782
 Papilio "Jamaica"



candidus
K.S. Brown
1976
Ecuador



chocoensis
K.S. Brown & W.W. Benson
1975
Colombia



hewitsoni [species] W.C. Hewitson
1875
 Heliconia Panama



leuce
E. Doubleday
1847
 Heliconia "Brazil"

38 sara

J.C. Fabricius
1793
 Papilio



sara
J.C. Fabricius
1793
 Papilio "Guyana"



apseudes
J. Hübner
1813
 Nerëis Brazil



brevimaculata
H.H. Riffarth
1901
Colombia



elektra
W.M. Neukirchen
1998
Colombia



fulgidus
H.F.E.J. Stichel
1906
Costa Rica



magdalena
H.W. Bates
1864
Colombia; Panama



sprucei
H.W. Bates
1864
Ecuador



theudela
W.C. Hewitson
1874
 Heliconia Panama



veraepacis
H.W. Bates
1864
Guatemala



williami
W.M. Neukirchen
1994
Trinidad

39 telesiphe

E. Doubleday
1847
 Heliconia



telesiphe
E. Doubleday
1847
 Heliconia Bolivia



cretacea
H. Neustetter
1916
"French Guiana"



sotericus
O. Salvin
1871
Ecuador

40 timareta

W.C. Hewitson
1867
 Heliconia



timareta
W.C. Hewitson
1867
 Heliconia Ecuador



florencia
N. Giraldo et al.
2008
Colombia



timoratus
G. Lamas
1998
Peru



[n. ssp.]
J. Mavarez et al.
MS
Peru



[n. ssp.]
K. Willmott
MS
Ecuador

41 tristero

A.V.Z. Brower
1996
Colombia

42 wallacei

T. Reakirt
1866




wallacei
T. Reakirt
1866
[Brazil]



araguaia
K.S. Brown
1976
Brazil (Go)



colon
G. Weymer
1891
[Brazil (Am)]



flavescens
G. Weymer
1891
Ecuador



kayei
H. Neustetter
1929
Trinidad



mimulinus
A.G. Butler
1873
Colombia

43 xanthocles

H.W. Bates
1862




xanthocles
H.W. Bates
1862
Guyana; "French Guiana"



buechei
W.M. Neukirchen
1992
Venezuela



cleoxanthe
H. Holzinger & R. Holzinger
1972
Venezuela



donatia
H. Fruhstorfer
1910
Brazil (MT)



explicata
K.S. Brown
1976
Colombia



hippocrene
H. Holzinger & K.S. Brown
1982
Bolivia



melete
C. Felder & R. Felder
1865
Colombia



melior
O. Staudinger
1897
Peru



melittus
O. Staudinger
1897
Peru



napoensis
H. Holzinger & K.S. Brown
1982
Ecuador



paraplesius
H.W. Bates
1867
Brazil (Ma)



quindecim
G. Lamas
1976
Peru



rindgei
H. Holzinger & K.S. Brown
1982
Colombia



similatus
J. Zikán
1937
Brazil (Am)



vala
O. Staudinger
1885
French Guiana



zamora
H. Holzinger & K.S. Brown
1982
Ecuador